England have spent several weeks training with portable hypoxic generators — colloquially dubbed 'Bane masks' by the squad — to prepare for Saturday's Nations Championship opener against the Springboks at Ellis Park.

Strength and conditioning coach Nathan Beardsley ran the players through high-intensity watt bike sessions at simulated altitudes of around 3,500 metres, roughly double Ellis Park's 1,753 metres above sea level. The approach was not primarily aimed at boosting red blood cell counts, but at conditioning the brain to manage acute breathlessness and preserving repeated sprint power under oxygen restriction.

"The mask works to sharpen the repeated sprint power and bank the breathless feeling. You're not actually changing the physiology a huge amount, but you're preparing that breathless feeling and then sharpening the repeated sprint effect, which programmes their muscle fibres to work at that fast pace," Beardsley explained.

England have also drilled the squad in structured recovery breathwork — diaphragmatic breathing and 'breath dumps' designed to clear carbon dioxide rapidly during lineout huddles, defensive sets or kicking windows. Beardsley flagged the mental dimension as equally critical, noting that the psychological weight of breathlessness, compounded by the occasion, is frequently what undoes visiting sides before the physical toll takes hold.

England's preparation has been shaped in part by a compressed schedule: after Saturday's Test at Ellis Park they fly back to the UK to face Fiji in Liverpool the following weekend before heading to Argentina to meet the Pumas.